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2011; Frac et al., 2018), including nutrient cycling (Gui et al., 2017) and the decomposition (Zak et al., 2019) and stabilisation of soil organic matter (Clemmensen et al., 2013). Moreover, through their many complex interactions with plant roots, fungi aid plant nutrient acquisition (Averill et al., 2019), pathogen (Marx, 1972) and drought resistance (Jayne & Quigley, 2014), and play a key role in shaping plant productivity and community dynamics (Mommer et al., 2018; Liang et al., 2020). Given this, understanding the key determinants of the diversity and composition of rhizosphere fungal communities is recognised as an important goal in terrestrial ecology (Singh et al., 2004; Francioli et al., 2020).