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A new neotropical genus, , with a single species, , is described from the Cordillera del Cóndor and Amazonian regions of southeastern Ecuador and is placed in the family Gesneriaceae, tribe Beslerieae. The placement of in the tribe Beslerieae is strongly supported by molecular sequence data generated from analyses of nuclear ribosomal DNA internal transcribed spacer region (ITS), and the chloroplast DNA intron, intergenic spacer region. Sequence data were generated for 33 species representing most of the generic diversity from the tribes Beslerieae and Napeantheae. is characterized by the following unique combination of relatively uncommon characters in the Gesneriaceae: arborescent habit; opposite leaf arrangement sometimes interrupted by alternate leaves; lepidote trichomes on vegetative and floral structures; small white flowers with a dorsal gibbosity at the base of the corolla tube; and a bivalved …