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This study examines the complex role of family networks in shaping adult psychological well-being over time. We examine the unique and interactive longitudinal influences of family structure (ie, composition and size) and negative family relationship quality on psychological well-being among young (ages 18–34), middle-aged (ages 35–49), and older adults (ages 50+). A sample of 881 adults (72% White; 26% Black) was drawn from the longitudinal Social Relations, Age, and Health Study. Structural equation modeling indicated that among young and middle-aged adults, increasing family negativity was associated with increases in depressive symptoms over time. In contrast, among older adults, lowered proportion of family in network and an increasing number of family members in the network (ie, family size) were associated with decreases in depressive symptoms. These findings were moderated by family …