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The vestimentiferan tubeworm Riftia pachyptila (Polychaeta: Sibloglinidae) often dominates early succession stages and high productivity habitats at low-temperature hydrothermal vents on the East Pacific Rise. We collected 8 aggregations of R. pachyptila and the associated epifaunal community at 2 discrete sites of diffuse hydrothermal activity, in December 2001 and December 2002. Because of the high spatial and temporal variability of the biotic and abiotic factors related to hydrothermal vent activity, significant differences in the structure and the composition of the community were expected to occur at the scale of either 1 yr or 500 m distance between very different sites. There was no significant difference in the temperature ranges of the diffuse flow between sites or years, even though the environmental conditions were very different at the 2 sites. At 1 site (Riftia Field), the diffuse hydrothermal fluids had …