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We distinguish two types of large multipolar cells designated sustained (C_s) and onset (O_c) choppers in the anterior posteroventral cochlear nucleus (A‐PVCN)/nerve root region on the basis of certain anatomical and physiological features. C_s axons head into the trapezoid body, while O_c axons use the intermediate acoustic stria of Held. At the electron microscopic (EM) level, collateral terminals of O_c axons contain pleomorphic vesicles; C_s terminals contain small round vesicles. C_s dendritic trees tend to be distributed in a stellate fashion while O_c dendritic trees tend to be elongated. At the EM level the sustained chopper somata are sparsely innervated while the proximal dendritic tree receives considerably more input. The O_c somata are highly innervated and this heavy innervation continues out onto the proximal dendrites. Distally the dendritic innervation falls off considerably for both categories. Physiologically …