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FIGURE 1. Automatic detection of the bone center using a section of a radius of the Upper Triassic (Carnian) stereospondyl Buettneria. Note that the program can also handle bone sections exhibiting color heterogeneity. most abrupt change in compactness is observed. This relative distance is the ratio between the distance to the center and the radius. This generally corresponds with the position of the transition zone between the cortical compacta and the medullary cavity (or spongiosa). S is the reciprocal of the slope of the compactness change at point P. It usually reflects the width of the transition zone between the cortical compacta and the medullary cavity (or the medullary spongiosa). Max is the upper asymptote. It generally reflects the compactness in the outermost cortex. Min is the lower asymptote. It generally reflects the compactness in the center of the medulla.