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All organisms require metal ions to complete their life cycle. Excess or shortage of essential metal ions is toxic to plants. Also, some heavy metals are toxic at all concentrations and hinder the functioning of plants. Therefore, plants including microbes have evolved metal homeostatic machineries to tackle toxic levels of metals inside the cell. Since a long time, scientists have investigated metal homeostasis mechanisms in plants. In last few decades, anthropogenic activities together with natural catastrophic events have increased the bioavailable concentration of heavy metals in the biosphere. Heavy metals are persistent in nature and cannot be biodegraded. Thus, heavy-metal pollution is becoming a threat to environment, agriculture, and human health. The microbes are the most sensitive creature to metal stress than the rest of soil fauna. Some plant-microbe interactions are beneficial under stress induced …