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Method–Image Analysis section. We have clarified that segments needed to be fully impregnated by Golgi stain to be included for analysis. Method–Image analysis section. We have clarified that branched spines were only counted as one spine. Method–Image analysis section. We have clarified how our image processing method ensured spines deeper in the tissue section were not missed from being counted. Method–Image analysis section. We have clarified that a 0.2µm step size was used to collect image stacks. Method–Image analysis section. We have clarified that spines above or below the dendrite with respect to the field of view are not visible with Golgi stain and hence no attempt to count or correct for them was made. Method–Golgi staining. We have now clarified that sections were sliced in the coronal plane. Method–Image analysis. We have clarified that CA1 basal dendritic segments were imaged …