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Forest soil organic matter (SOM) is derived from both above- and belowground plant inputs, with a significant portion of this plant-derived organic carbon being processed by fungal communities prior to becoming SOM. Despite the central role of fungi in the processing of litter and root inputs, little is known about the relationship between fungal community composition and the relative conversion of aboveground litter versus root inputs into SOM. The Detrital Input and Removal Treatment (DIRT) experiments offer a system in which to test such questions, wherein aboveground litter and root inputs to forest soils are chronically excluded independently and in combination. In this study, we characterized fungal biomass (ergosterol), community composition (ITS2 metabarcoding), and community assembly (via null modeling) under root and litter exclusion treatments in a northern temperate forest DIRT experiment, and we …