The Yellowstone hotspot commonly is thought to result from a stationary mantle plume rooted in the lower mantle over which North America moves. Yet Yellowstone’s initiation and its association with the “backward” propagating Newberry hotspot across eastern Oregon pose difficult questions to those explaining Yellowstone as a simple consequence of a deep-seated plume. Teleseismic investigations across the Yellowstone topographic swell reveal:(1) the swell is held up by buoyant mantle of two types—partially molten mantle (of low seismic velocity) beneath the hotspot track and basalt-depleted mantle (of high velocity) beneath the rest of the swell;(2) an upwarped 660 km discontinuity beneath the Yellowstone hotspot track, as expected for relatively hot mantle at that depth, and an upwarped 410 km discontinuity, indicative of relatively cool mantle at this depth; and (3) anisotropic mantle with a preferred northeast orientation of olivine a axis, consistent with the strain expected for both plate motion and hotspot asthenosphere flow. Imaged mantle velocities can be reconciled with a plume hypothesis only if melt buoyancy within the hotspot asthenosphere drives convection, with melt segregating from the mantle beneath Yellowstone and residuum being deposited adjacent to the upwelling. Once such convection is admitted, an alternative, nonplume explanation for Yellowstone is possible, which has propagating convective rolls organized by the sense of shear across the asthenosphere. This explanation has the appeal that expected asthenospheric shear beneath the northwest United States predicts both the Yellowstone and Newberry hotspots with a single (upper mantle) process.