The incidence of predation by largemouth bass Micropterus salmoides on fish in Lake Conroe, Texas, was examined over 7 years at two disparate levels of habitat complexity. When areal coverage of submersed vegetation ranged from 39 to 44% of the reservoir's 8,100 hectares, largemouth bass 100 mm and smaller in total length consumed fish infrequently; most did not consume fish until they reached lengths of 140 mm and more, Following the elimination of all submersed vegetation by grass carp Ctenopharyngodon idella, fish were consumed by most largemouth bass 60 mm or longer. The initiation of piscivory at smaller sizes resulted in significantly faster first‐year growth for all largemouth bass year‐classes produced after vegetation removal. Although shifts in the structure of the forage fish community occurred, ample fish prey existed for largemouth bass before and after vegetation removal. The onset of piscivory remained similar for largemouth bass collected along the dam riprap, where vegetation was absent throughout the study. These observations support the hypothesis that habitat complexity, as mediated by vegetation abundance, was the principal factor regulating piscivory by largemouth bass in the littoral zone of Lake Conroe.